BSc (hons) in Zooloy, University of Birmingham UK 1947 PhD in Zoology (Embryology), University of London 1951, supervised by Sir Gavin deBeer and Michael Abercrombie. Apart from brief periods in Anatomy Department, Cambridge and Department of Biology, St Bartholomew's Hospital Medical School, academic posts have all been at UCL (University College London). Retired 1991 as Emeritus Professor in Embryology. Research predominantly on early chick embryos concerned mainly with problems of tissue interaction, gastrulation and segmentation. Founder member of Embryologists' Club, the predecessor of BSDB (British Society of Developmental Biology). Served on several editorial boards, including JEEM… (now Development) and Anatomy and Embryology. Publications: approximately 120, including 2 books, 5 edited books.